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nutrient deficient?

Here are some images of my young Red Paper Lanterns, what does the darker color within the leaves indicate?
 
Are the plants deficient in a particular nutrient?
 
rpl1.jpg

 
 
rpl2.jpg

 
 
rpl3.jpg
 
Yes, all 6 of them.  4 Red Lanterns and 2 Brazilian Starfish.
 
12 hours of 6000 Kelvin light and at least 2 to 3 hours of sunlight each day.
 
 
 
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